Current position in this corpus

A major trend across this collection is the move away from simple epithelial balls toward organoids with multiple coordinated lineages, appendages, or anatomical boundary relationships.

Strong supporting sources

Working synthesis

  • Lancaster, Lee, and some cardiac papers show the value of emergent tissue complexity.
  • Koike and Dardano explicitly pursue linked developmental programs across neighboring organs or hematopoietic compartments.
  • Olijnik and Matkovic Leko push organoid logic into niche or disease contexts where multicellular composition is central to the model.

Main tension

  • biological richness versus protocol simplicity and scalability
  • compositional realism versus clean mechanistic interpretation

Open questions

  • Which extra tissue compartments materially change the answer to the biological question?
  • When does added complexity improve realism, and when does it mostly increase variability?